You are right. I didn't even realised that the male of G. lebadea have no spots at all.
I couldn't ID to species because I realised except for M. aria, all the rest (Matapa) found in Borneo have orange/yellow tornal cilia.
The Cepora is confirmed to be C. pactolicus. Check http://imgl-a.dena.ne.jp/exl3/201011...51570729_1.jpg
TL Seow![]()